Influences on the feeding behavior of three mammals in the Maruyama Zoo: bears, elephants and chimpanzees

  • Published source details Morimura N. & Ueno Y. (1999) Influences on the feeding behavior of three mammals in the Maruyama Zoo: bears, elephants and chimpanzees. Journal of Applied Animal Welfare Science, 2, 169-186.


This study is summarised as evidence for the following.

Action Category

Primates: Change the number of feeds per day

Action Link
Management of Captive Animals
  1. Primates: Change the number of feeds per day

    A before-and-after study in 1999 in Japan (Morimura & Ueno 1999) found that when food was given twice a day instead of once a day to a group of chimpanzees Pan troglodytes the amount of time spent feeding increased. On average, feeding time contributed to 7% of the behaviours expressed when one feed was offered per day and 24% when two feeds were offered per day. A group of five chimpanzees was observed for 15 days over five one hour time periods when all food was provided in the afternoon at 15:00 h and when one feed was given between 10:00 h and 11:00 h and a second feed at 15:00 h.    (CJ)

Output references
What Works 2021 cover

What Works in Conservation

What Works in Conservation provides expert assessments of the effectiveness of actions, based on summarised evidence, in synopses. Subjects covered so far include amphibians, birds, mammals, forests, peatland and control of freshwater invasive species. More are in progress.

More about What Works in Conservation

Download free PDF or purchase
The Conservation Evidence Journal

The Conservation Evidence Journal

An online, free to publish in, open-access journal publishing results from research and projects that test the effectiveness of conservation actions.

Read the latest volume: Volume 21

Go to the CE Journal

Discover more on our blog

Our blog contains the latest news and updates from the Conservation Evidence team, the Conservation Evidence Journal, and our global partners in evidence-based conservation.

Who uses Conservation Evidence?

Meet some of the evidence champions

Endangered Landscape ProgrammeRed List Champion - Arc Kent Wildlife Trust The Rufford Foundation Save the Frogs - Ghana Mauritian Wildlife Supporting Conservation Leaders
Sustainability Dashboard National Biodiversity Network Frog Life The international journey of Conservation - Oryx Cool Farm Alliance UNEP AWFA Bat Conservation InternationalPeople trust for endangered species Vincet Wildlife Trust