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Providing evidence to improve practice

Individual study: Are species listed in the Annex II of the Habitats Directive better represented in Natura 2000 network than the remaining species? A test using Spanish bats

Published source details

Lisón F., Sánchez-Fernández D. & Calvo J.F. (2015) Are species listed in the Annex II of the Habitats Directive better represented in Natura 2000 network than the remaining species? A test using Spanish bats. Biodiversity and Conservation, 24, 2459-2473


This study is summarised as evidence for the intervention(s) shown on the right. The icon shows which synopsis it is relevant to.

Legally protect bat habitats Bat Conservation

A study in 2015 of protected areas in Spain (Lisón et al 2015) found that the distributions of 10 of 11 target bat species and 13 of 18 non-target bat species overlapped with protected ‘Special Conservation Areas’ (SACs) significantly more than expected by chance. The distributions of nine of 11 target bat species and 13 of 18 non-target bat species also overlapped with ‘Special Protection Areas’ (SPAs) designated to protect birds. The amount of overlap between bat species distributions and either of the protected area types did not differ significantly between target and non-target species. Both SPAs and SACs were part of the legally protected European Natura 2000 network. Target species were of highest conservation concern and listed in Annex II of the European Habitats Directive. All other (non-target) bat species were listed in Annex IV. The mean percentage overlap between species distributions (grid cells in which the species occurred) and the protected areas were calculated using an existing bat dataset for mainland Spain and the Balearic Islands.

(Summarised by Anna Berthinussen)